College of Resource and Environment, Jilin Agricultural University, Changchun, 130118, PR China. College of Plant Sciences, Jilin University, Changchun, 130062, China. Department of Computer Science, Iowa State University, Ames, IA, USA, 50011. College of Resource and Environment, Jilin Agricultural University, Changchun, 130118, PR China. Electronic address: haozhang100@163.com. School of Science and Technology, Georgia Gwinnett College, Lawrenceville, GA, USA, 30024. Electronic address: fsun@ggc.edu.